{"id":1627,"date":"2021-04-10T00:15:16","date_gmt":"2021-04-10T00:15:16","guid":{"rendered":"https:\/\/www.musicianscience.org\/?p=1627"},"modified":"2021-04-10T00:15:18","modified_gmt":"2021-04-10T00:15:18","slug":"brain-activity-in-musical-improvisers-revealed","status":"publish","type":"post","link":"https:\/\/www.musicianscience.org\/index.php\/brain-activity-in-musical-improvisers-revealed\/","title":{"rendered":"Brain activity in musical improvisers revealed"},"content":{"rendered":"\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"250\" height=\"375\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-250px-Trevor_Pinnock_2.jpg\" alt=\"\" class=\"wp-image-1657\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-250px-Trevor_Pinnock_2.jpg 250w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-250px-Trevor_Pinnock_2-200x300.jpg 200w\" sizes=\"auto, (max-width: 250px) 100vw, 250px\" \/><\/figure><\/div>\n\n\n\n<p class=\"has-text-align-center\"><strong>Title of paper under discussion<\/strong><\/p>\n\n\n\n<p class=\"has-text-align-center\">Cortical Regions Involved in the Generation of Musical Structures during Improvisation in Pianists<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong>Authors<\/strong><\/p>\n\n\n\n<p class=\"has-text-align-center\">Sara L. Bengtsson, Mih\u00e1ly Cs\u00cdkszentmih\u00e1lyi and Fredrik Ull\u00e9n<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong>Journal<\/strong><\/p>\n\n\n\n<p class=\"has-text-align-center\">Journal of Cognitive Neuroscience, vol 19, no 5, pp. 830\u2013842<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong><a rel=\"noreferrer noopener\" href=\"http:\/\/www.brainmusic.org\/EducationalActivitiesFolder\/Bengtsson_improv2007.pdf\" target=\"_blank\">Link to paper<\/a> (free access)<\/strong><\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"276\" height=\"183\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-KarolUnknown.jpeg\" alt=\"\" class=\"wp-image-1662\"\/><figcaption><em><span class=\"has-inline-color has-vivid-purple-color\">Above &#8211; Karolinska Institute, Stockholm<\/span><\/em><\/figcaption><\/figure><\/div>\n\n\n\n<p class=\"has-text-align-center\"><strong>Overview<\/strong><\/p>\n\n\n\n<p><span class=\"has-inline-color has-vivid-red-color\">Scan the brain of someone in the act of selecting freely, for example moving a hand hand at will or choosing any number from 1 to 10, and a couple of regions will typically \u201clight up\u201d: the \u2018dorsolateral prefrontal cortex\u2019 and \u2018rostral premotor cortex\u2019.  Would this be the same for an improvising musician?<\/span><\/p>\n\n\n\n<p><span class=\"has-inline-color has-vivid-red-color\">Using functional magnetic resonance imaging (fMRI), Bengtsson and her colleagues scanned the brains of 11 professional pianists as they improvised on a simple prescribed melody, a task called \u2018Improvise\u2019.  The pianists were then asked to repeat that same improvisation as accurately as possible from memory, a task called \u2018Reproduce\u2019.  Finally the pianist was asked to do a free improvisation, with no need to memorise it, a task called \u2018FreeImp\u2019.<\/span><\/p>\n\n\n\n<p class=\"has-text-color has-vivid-red-color\">The scientists reasoned that by contrasting the \u2018Improvise\u2019 scan with the \u2018Reproduce\u2019 one they might reveal those regions of brain that were active due to improvisation, as the contrast would delete brain activity present in <em>both<\/em> tasks (eg pianistic movements and sensations).  In case the \u2018Improvise\u2019 scans were also picking up \u2018memorising\u2019 brain activity needed for that task, the researchers compared their \u2018Improvise vs Reproduce\u2019 contrasts with \u2018FreeImp vs Resting state\u2019 contrasts.   Brain areas active in both contrasts, they argued, must be underpinning the act of musical improvisation.<\/p>\n\n\n\n<p class=\"has-text-color has-vivid-red-color\">As predicted, such activated brain regions did include the right \u2018dorsolateral prefrontal cortex\u2019 and \u2018rostral premotor areas\u2019, plus the posterior part of the left \u2018superior temporal gyrus\u2019.  The authors suggest these regions \u201care part of a network involved in musical creation\u201d.<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong>Method<\/strong><\/p>\n\n\n\n<p>Eleven Swedish male right-handed concert pianists volunteered for the study.  Though lying in an fMRI scanner throughout the experiment, each participant was able to \u2018play the piano\u2019 (a one-octave piano midi keyboard)  and read music (from a screen via a binocular\/mirror system).<\/p>\n\n\n\n<p>The experiment was composed of four parts, or \u2018conditions\u2019: Improvise, Reproduce, Rest and FreeImp.  In all conditions a musical score consisting of a simple eight-bar musical melody was displayed, for example:<\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"515\" height=\"208\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-1a.png\" alt=\"\" class=\"wp-image-1646\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-1a.png 515w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-1a-300x121.png 300w\" sizes=\"auto, (max-width: 515px) 100vw, 515px\" \/><\/figure><\/div>\n\n\n\n<p>After 8 seconds of silently reading the score the participant was given instructions according to the experimental condition.<\/p>\n\n\n\n<p>For Improvise, they were asked to use the displayed melody as a basis for an improvisation. The instructions to the participants were to \u201cemploy any kind of modifications of the presented melodic template they wished [\u2026]They were also instructed to memorize the performance for subsequent reproduction\u201d<\/p>\n\n\n\n<p>For Reproduce, again using the keyboard, \u201cthe task was to reproduce, as faithfully as possible, the improvisation previously made upon the same melody.\u201d<\/p>\n\n\n\n<p>For Rest, the instructions were to remain \u201crelaxed, viewing the screen without any movements.\u201d<\/p>\n\n\n\n<p>For FreeImp, \u201cthe instruction was to improvise on the melody, as in Improvise, but without trying to memorize the performance.\u201d<\/p>\n\n\n\n<p>The experiment was repeated 9 times, with a different melody used each time.<\/p>\n\n\n\n<p>As well as constantly monitoring each participant\u2019s brain activity, the experimenters also made a careful midi-recording of each of their piano performances; this enabled them, for example, to record how accurately the pianist had managed to reproduce a given \u2018Improvise\u2019 improvisation in the subsequent Reproduce condition, and to compare FreeImp complexity with Improvise complexity.<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong>Results<\/strong><\/p>\n\n\n\n<p><em>Styles of improvisation<\/em><\/p>\n\n\n\n<p>Looking across all the improvisation recordings, Bengtsson and her colleagues counted up 11 different types of musical device used by the pianists to elaborate or modify their given melodies (or \u2018templates\u2019): <\/p>\n\n\n\n<p>\u201ci) Insertion of a fast group of one or more grace notes before a template note; (ii) substitution of a template note for another note; (iii) figuration, that is, expansion of the original template into melodic figures; (iv) insertion of a trill on a template note; (v) filling in, that is, insertion of chromatic or diatonic scales between template notes; (vi) repetitions of template notes; (vii) elimination of template notes; (viii) insertion of figures giving a broken two-part polyphony; (ix) rhythmization of the template; (x) insertion of a tremolo, that is, a trill-like figure between two notes with a larger interval than a second; and finally, (xi) switching of tonality from major to minor.\u201d  Examples of (i) to (v) are shown here:<\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"577\" height=\"610\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-1b.png\" alt=\"\" class=\"wp-image-1647\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-1b.png 577w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-1b-284x300.png 284w\" sizes=\"auto, (max-width: 577px) 100vw, 577px\" \/><\/figure><\/div>\n\n\n\n<p><em>Accuracy of the Reproduce performances<\/em><\/p>\n\n\n\n<p>To give their fMRI analysis credence, the scientists needed to establish whether the Reproduce performances were similar enough in execution to the Improvise performances they were supposed to mimic.  If so, then their proposal &#8211; that any contrast between an Improvise scan and its respective Reproduce scan would reveal the brain activity involved in the act of improvisation &#8211;  should be sound.  If not, then such a contrast may also be showing up brain activity due to the differences in physical execution between the two types of performance.<\/p>\n\n\n\n<p>Comparing Improvise performances with Reproduce performances,  the scientists found there was no significant difference in the number of notes played, nor in performance duration.   To further investigate how well each improvisation was reproduced the researchers carried out a note-by-note comparison between each Improvise performance and its subsequent Reproduce performance, assigning each comparison an accuracy score called \u2018LED\u2019 (\u2018Levenshtein edit distance\u2019).  They found that \u201cthe participants were able to reproduce their performances in Improvise with remarkable accuracy in the Reproduce condition.\u201d  <\/p>\n\n\n\n<p><em>FMRI scan contrasts<\/em><\/p>\n\n\n\n<p>Having established that the pianists were remarkably accurate in their Reproduce performances, the scientists could move on to the analysis of brain imagery confident that the Improvise-Reproduce contrasts would indeed reveal brain activity solely involved in the act of improvisation (though perhaps also including brain activity associated with the act of memorising).<\/p>\n\n\n\n<p>Four brain regions were identified as being more active during Improvise than Reproduce: the right Dorsolateral Prefrontal Cortex (DLPFC), Rostral Premotor Cortex (RPC), left Superior Temporal Gyrus (STG) and the middle occipital gyri (MOG):<\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"507\" height=\"388\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-4.png\" alt=\"\" class=\"wp-image-1649\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-4.png 507w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-4-300x230.png 300w\" sizes=\"auto, (max-width: 507px) 100vw, 507px\" \/><figcaption><em><span class=\"has-inline-color has-vivid-purple-color\">Above &#8211; some areas of the brain active during improvisation: right DLPFC (dorsolateral prefrontal cortex), left STG (superior temporal gyrus) and PMD (dorsal premotor codex, part of the rostral premotor cortex. or RPC) <\/span><\/em><\/figcaption><\/figure>\n\n\n\n<p>in order to establish that these brain regions were \u2018lighting up\u2019 due to the act of improvisation and not memorising, the researchers also contrasted the scans taken during the FreeImp condition with those of the Rest condition.  They reasoned that any areas active in the FreeImp-Rest contrast that were <em>also<\/em> active in the Improvise-Reproduce contrast <em>must<\/em> be solely involved in improvisation, and not also memorisation, because during FreeImp the instruction was not to memorise the performance.<\/p>\n\n\n\n<p>It turned out that the all these four brain regions, or areas very close to them, were also active in the FreeImp-Rest contrast, leading the researchers to assert with more confidence that the right DLPFC, the right and left RPC, the left STG and the right and left MOG are all critical for the act of musical improvisation. In the words of the authors \u201cwe therefore argue that the activity in these brain regions reflects neural processes involved in the generation of new musical material during improvisation.\u201d<\/p>\n\n\n\n<p>Another piece of analysis further implicated a specific sub-region of RPC, the right pre-Supplementary Motor Area (right pre-SMA), as being especially involved in the act of improvisation.   By comparing the <em>complexity<\/em> of each Improvisation with that performer\u2019s right pre-SMA brain activity during its execution, the scientists discovered that the more complex the improvisation, the more active was the right pre-SMA.  In other words, the harder the pianist was improvising, the harder the right pre-SMA was working.<\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"892\" height=\"364\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-5.png\" alt=\"\" class=\"wp-image-1652\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-5.png 892w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-5-300x122.png 300w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-fig-5-768x313.png 768w\" sizes=\"auto, (max-width: 892px) 100vw, 892px\" \/><figcaption><em><span class=\"has-inline-color has-vivid-purple-color\">Above &#8211; the greater the complexity of the improvisation (x-axis) the more the pre-Supplementary Motor Area (pre-SMA) becomes active (y-axis)<\/span><\/em><\/figcaption><\/figure>\n\n\n\n<p>In a similar vein, the DLPFC was generally more active during FreeImp than Improvise, which could be related to the slightly higher level of improvisational complexity seen in Freeimp compared with Improvise.<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong>Discussion<\/strong><\/p>\n\n\n\n<p>What is it about these particular brains regions that makes them \u2018light up\u2019 during improvisations?<\/p>\n\n\n\n<p>The authors looked at the findings of other brain researchers in an effort to ascribe specific improvisational roles to each of these regions.<\/p>\n\n\n\n<p><em>Dorsolateral Prefrontal Cortex (DLPFC)<\/em><\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Brodmann_area_46_animation_small.gif\" alt=\"\" class=\"wp-image-1653\"\/><figcaption><em><span class=\"has-inline-color has-vivid-purple-color\">Above &#8211; Dorsolateral prefrontal cortex<\/span><\/em> (gif from <a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Brodmann_area_46_animation_small.gif\" target=\"_blank\" rel=\"noreferrer noopener\">BodyParts3D<\/a>)<\/figcaption><\/figure><\/div>\n\n\n\n<p>Previous research papers on many different types of <em>motor task<\/em> have consistently implicated the DLPFC when movement is freely chosen compared with movement in  \u2018repetitive or externally determined\u201d tasks. Similarly, the authors explained, DLPFC acitivity is related to free selection in <em>cognitive tasks<\/em>, such as word generation or number generation.<\/p>\n\n\n\n<p>Some research suggests that the DLPFC is especially involved in <em>paying attention<\/em> to free choice selection, while other research implicates it in the actual selection process itself.  Memory researchers have suggested that the DLPFC is implicated in inhibiting stereotyped, habitual responses in favour of random (or at least pseudo-random) ones, achieving this by constantly monitoring such responses in \u2018working memory\u2019.<\/p>\n\n\n\n<p>In addition to these specific roles &#8211; attention to action, monitoring in working memory, response selection and suppression of stereotype responses &#8211; the authors claim that their results are consistent with \u201ca vast body of neurological and experimental data\u201d claiming a \u201ccentral role for the DLPFC in planning and performance of novel or complex behavioural sequences, including language and thought\u201d.  Thus, claims Bengtsson, the DLPFC has \u201cone central function\u201d during improvisation, which is to be \u201csupervisory: to maintain and execute an overall plan for the improvisation through top-down influences on the activity\u201d, with the RPC areas &#8211; with which it has rich connections &#8211; subordinate to its control.  In this way \u201ca whole set of freely selected modifications of the original melody\u201d can be organised in time \u201caccording to a musically meaningful overall plan\u201d.  <\/p>\n\n\n\n<p><em>Rostral premotor cortex (RPC)<\/em><\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"600\" height=\"600\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Brodmann_area_6_lateral.jpg\" alt=\"\" class=\"wp-image-1654\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Brodmann_area_6_lateral.jpg 600w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Brodmann_area_6_lateral-300x300.jpg 300w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Brodmann_area_6_lateral-150x150.jpg 150w\" sizes=\"auto, (max-width: 600px) 100vw, 600px\" \/><figcaption><span class=\"has-inline-color has-vivid-purple-color\"><i>Above &#8211; premotor cortex and supplementary motor area; the front of this strip is the rostral premotor cortex (RPC)<\/i><\/span>, <em><span class=\"has-inline-color has-vivid-purple-color\">active during improvisation<\/span><\/em> <em><span class=\"has-inline-color has-vivid-purple-color\">[image from <a rel=\"noreferrer noopener\" href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Brodmann_area_6_lateral.jpg\" target=\"_blank\">BodyParts 3D<\/a>]<\/span><\/em><\/figcaption><\/figure>\n\n\n\n<p>Having suggested that the RPC is subordinate to the DLPFC in the act of improvisation, our authors identify two sub-areas of RPC &#8211; the pre-Supplementary Motor Cortex (pre-SMA) and the Dorsal Premotor Cortex (PMD) &#8211; that may each support different improvisatory roles.<\/p>\n\n\n\n<p>The pre-SMA, according to previous research, is especially involved in timing and rhythmic sequence performance, leading our authors to \u201ctherefore suggest that the pre-SMA activity during improvisation may be particularly related to decisions about timing and rhythmic patterning.\u201d  The PMD, on the other hand, has been implicated in the transformation of information held in \u2018working memory\u2019 into a sequence of movements in the correct order.<\/p>\n\n\n\n<p>Thus, claim the authors, the role of the RPC in improvisation could be the timing (pre-SMA) and sequencing (PMD) of movements, such that the PMD is more involved with <em>melodic<\/em> structures and the pre-SMA with <em>rhythmic<\/em> structures.<\/p>\n\n\n\n<p><em>Superior Temporal Gyri (STG)<\/em><\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"900\" height=\"900\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Cerebrum_-_superior_temporal_gyrus_-_lateral.png\" alt=\"\" class=\"wp-image-1658\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Cerebrum_-_superior_temporal_gyrus_-_lateral.png 900w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Cerebrum_-_superior_temporal_gyrus_-_lateral-300x300.png 300w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Cerebrum_-_superior_temporal_gyrus_-_lateral-150x150.png 150w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/impro-Cerebrum_-_superior_temporal_gyrus_-_lateral-768x768.png 768w\" sizes=\"auto, (max-width: 900px) 100vw, 900px\" \/><figcaption><span class=\"has-inline-color has-vivid-purple-color\"><i>Above &#8211; Superior Temporal Gyrus, the posterior part of which is active during improvisation [image from <\/i><a rel=\"noreferrer noopener\" style=\"font-style: italic;\" href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Cerebrum_-_superior_temporal_gyrus_-_lateral.png\" target=\"_blank\">BodyParts 3D<\/a><\/span>]<\/figcaption><\/figure><\/div>\n\n\n\n<p>The posterior STG is a brain region that \u201chas consistently been found active in studies that require auditory\u2013motor integration, such as rhythmic sequence performance[\u2026] and vocal rehearsal of words or music\u201d.  Bengtsson suggests that its increased activity evident in her pianists may reflect the increased reliance on \u201cauditory-motor feedback loops\u201d demanded by improvisation, \u201cfor example to adapt the improvisation to ongoing performance.\u201d.  The same region has also been shown to be involved in \u201cauditory working memory\u201d, and \u201cthus may be of importance for auditory monitoring of the ongoing performance\u201d.   The posterior STG also has a role in word retrieval in linguistic tasks &#8211; perhaps, suggest the authors, it is similarly involved in the retrieval of grace notes, trills etc from a \u201ccommon \u201cvocabulary\u201d of ornaments used in Western art music\u201d<\/p>\n\n\n\n<p><em>Middle occipital gyrus (MOG)<\/em><\/p>\n\n\n\n<div class=\"wp-block-image\"><figure class=\"aligncenter size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"1024\" src=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/improv1024px-Gyri_of_lateral_cortex.png\" alt=\"\" class=\"wp-image-1659\" srcset=\"https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/improv1024px-Gyri_of_lateral_cortex.png 1024w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/improv1024px-Gyri_of_lateral_cortex-300x300.png 300w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/improv1024px-Gyri_of_lateral_cortex-150x150.png 150w, https:\/\/www.musicianscience.org\/wp-content\/uploads\/2021\/04\/improv1024px-Gyri_of_lateral_cortex-768x768.png 768w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption><em><span class=\"has-inline-color has-vivid-purple-color\">Above &#8211; Middle Occipital Gyrus (MOG), coloured blue and labelled at the lower right hand side of the diagram (courtesy of <a rel=\"noreferrer noopener\" href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Gyri_of_lateral_cortex.png\" target=\"_blank\">Frank Gaillard<\/a>) <\/span><\/em><\/figcaption><\/figure><\/div>\n\n\n\n<p>Heightened activity in this brain area, known to process higher-order visual input including reading music, is thought by the authors to \u201cbe due to a more intense visual processing of the musical score when this was used as a basis for improvisations, rather than as a template to recall a previous performance.\u201d<\/p>\n\n\n\n<p>In conclusion the authors declare \u201cWe have provided evidence that a set of frontal and temporal association areas [DLPFC, RPC and STG] are specifically involved in the free creation of musical structure during improvisation.\u201d<\/p>\n\n\n\n<p class=\"has-text-align-center\"><strong>Coda<\/strong><\/p>\n\n\n\n<p class=\"has-text-align-center\"><a href=\"https:\/\/www.youtube.com\/watch?v=jFnM9C8jCdE\" target=\"_blank\" rel=\"noreferrer noopener\">Improvised Passacaglia in D minor<\/a><\/p>\n\n\n\n<p class=\"has-text-align-center\">Alexander Weimann &#8211; harpsichord<\/p>\n","protected":false},"excerpt":{"rendered":"<p>Title of paper under discussion Cortical Regions Involved in the Generation of Musical Structures during Improvisation in Pianists Authors Sara L. Bengtsson, Mih\u00e1ly Cs\u00cdkszentmih\u00e1lyi and Fredrik Ull\u00e9n Journal Journal of Cognitive Neuroscience, vol 19, no 5, pp. 830\u2013842 Link to paper (free access) Overview Scan the brain of someone in the act of selecting freely, &hellip;<\/p>\n<p class=\"read-more\"> <a class=\"\" href=\"https:\/\/www.musicianscience.org\/index.php\/brain-activity-in-musical-improvisers-revealed\/\"> <span class=\"screen-reader-text\">Brain activity in musical improvisers revealed<\/span> Read More \u00bb<\/a><\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"jetpack_post_was_ever_published":false,"_jetpack_newsletter_access":"","_jetpack_dont_email_post_to_subs":false,"_jetpack_newsletter_tier_id":0,"_jetpack_memberships_contains_paywalled_content":false,"_jetpack_memberships_contains_paid_content":false,"footnotes":""},"categories":[14,25],"tags":[],"class_list":["post-1627","post","type-post","status-publish","format-standard","hentry","category-creativity","category-musician"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v25.7 - 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